Population Demography of Northern Spotted Owls

Eric D. Forsman, who collaborated with a team of twenty-seven researchers to produce this report, is a Wildlife Biologist with the USDA Forest Service's Pacific Northwest Research Station, and also holds a courtesy faculty position as an Associate Professor in the Department of Fisheries and Wildlife at Oregon State University.

• INTRODUCTION STUDY AREAS FIELD METHODS ANALYTICAL METHODS DEVELOPMENT OF COVARIATES Barred Owl covariate Habitat covariates Weather and climate covariates Land ownership, ecoregion, and latitude covariates Reproduction covariate FECUNDITY Individual study areas Meta-analysis of fecundity APPARENT SURVIVAL Individual study areas Meta-analysis of apparent survival ANNUAL RATE OF POPULATION CHANGE Individual study areas Estimates of realized population change Meta-analysis of annual rate of population change STATISTICAL CONVENTIONS WORKSHOP PROTOCOLS RESULTS FECUNDITY Individual study areas Meta-analysis of fecundity APPARENT SURVIVAL Individual study areas Meta-analysis of apparent survival on all areas Meta-analysis of apparent survival on the eight NWFP monitoring areas Potential cost of reproduction on survival Effects of Barred Owls on recapture and survival ANNUAL RATE OF POPULATION CHANGE Individual study areas Estimates of realized population change Meta-analysis of annual rate of population change DISCUSSION FECUNDITY APPARENT SURVIVAL ANNUAL RATE OF POPULATION CHANGE AND REALIZED RATES OF POPULATION CHANGE Individual study areas Meta-analysis of annual rate of population change STATUS OF OWL POPULATIONS IN THE EIGHT NWFP MONITORING AREAS ASSOCIATIONS BETWEEN DEMOGRAPHIC PARAMETERS AND COVARIATES Cost of reproduction on survival Weather and climate Barred Owls Habitat POTENTIAL BIASES IN ESTIMATES OF DEMOGRAPHIC PARAMETERS Fecundity Apparent survival Annual rate of population change Estimating goodness-of-fit and overdispersion SUMMARY, CONCLUSIONS, and RECOMMENDATIONS ACKNOWLEDGMENTS LITERATURE CITED APPENDICES TABLES FIGURES

"One of the most exact and large-scale metapopulation analyses in the world." Luonnon Tutkija