Michael E. Hochberg is a Research Director with the Centre National de la Recherche Scientifique. A recipient of the Silver Medal of the CNRS in 1997, he employs both empirical and theoretical approaches toward understanding fundamental and applied aspects of evolution in host-parasite associations. Anthony R. Ives is an Associate Professor in the Departments of Zoology and Entomology at the University of Wisconsin-Madison. He is a theoretical ecologist who conducts empirical research on aphid pests and their parasitoids in agricultural systems.

• Preface xi List of Contributors xiii One: Introduction by Michael E. Hochberg and Anthony R. Ives 3 PART ONE: POPULATION DYNAMICS 15 Two: Host Location and Selection in the Field by Jgr6me Casas 17 Three: Effects of Parasitoid Clutch Size on Host-Parasitoid Population Dynamic by George E. Heimpel 27 Four: Host-Parasitoid Models: The Story of a Successful Failure by Carlos Bernstein 41 Five: A Field Guide to Studying Spatial Pattern Formation in Host-Parasitoid Systems by Susan Harrison 58 Parasitoid Spread: Lessons for and from Invasion Biology by Alan Hastings 70 Seven: Landscape Ecology of Parasitism by Jens Roland 83 PART TWO: POPULATION DIVERSITY 101 Eight: The Evolution of Parasitoid Egg Load by Minus van Baalen 103 Nine: Host Resistance, Parasitoid Virulence, and Population Dynamics by H. C. J. Godfray 121 Ten: Developmental Traits and Life-History Evolution in Parasitoids by Michael R. Strand 139 Eleven: Host Specificity and Trophic Relationships of Hyperparasitoids by Jacques Brodeur 163 Twelve: Comparing Parasitoid-Dominated Food Webs with Other Food Webs: Problems and Future Promises by Marcel Holyoak 184 Thirteen: Species Coexistence in Parasitoid Communities: Does Competition Matter? by Bradford A. Hawkins 198 PART THREE: POPULATION APPLICATIONS 215 Fourteen: Biological Control: The Need for Realistic Models and Experimental Approaches to Parasitoid Introductions by Nick Mills 217 Fifteen: Parasitoid Populations in the Agricultural Landscape by Teja Tscharntke 235 Sixteen: Threats, Flies, and Protocol Gaps: Can Evolutionary Ecology Save Biological Control? by Bernard D. Roitberg 254 Seventeen: "What, Conserve Parasitoids?" by Michael E. Hochberg 266 Eighteen: Conclusions: Debating Parasitoid Population Biology over the Next Twenty Years by Anthony R. Ives and Michael E. Hochberg 278 References 305 Index 359

"An excellent edited volume that provides a useful synthesis of knowledge in the field and clear guidance for future research goals. The book is not an exhaustive review of the field, but it is more readable because it is focused on a limited number of specific problems... [It] is concise and coherent enough so that readers will want to read the entire volume."--Timothy P. Craig, Ecology "A useful and interesting book."--Donald H. Feener, Jr.,The Quarterly Review of Biology